问错误信息："'NoneType‘对象不可调用“。-收到zappa部署的django在AWS Lambda上的应用程序

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{'message': 'An uncaught exception happened while servicing this request. You can investigate this with the `zappa tail` command.', 'traceback': ['Traceback (most recent call last):', ' File /var/task/handler.py, line 540, in handler with Response.from_app(self.wsgi_app, environ) as response:', ' File /var/task/werkzeug/wrappers/base_response.py, line 287, in from_app return cls(*_run_wsgi_app(app, environ, buffered))', ' File /var/task/werkzeug/wrappers/base_response.py, line 26, in _run_wsgi_app return _run_wsgi_app(*args)', ' File /var/task/werkzeug/test.py, line 1119, in run_wsgi_app app_rv = app(environ, start_response)', TypeError: 'NoneType' object is not callable]}

{
  "production": {
    "aws_region": "eu-west-2",
    "django_settings": "app.settings",
    "profile_name": "deployment",
    "project_name": "app",
    "runtime": "python3.6",
    "s3_bucket": "zappa-deployment-uploads",
    "slim_handler": true,
    "exclude": [".ebextensions/", ".elasticbeanstalk/", "webpack/", "app/static/"],
    "vpc_config" : {
    "SubnetIds": [ "..."],
    "SecurityGroupIds": ["..."]
  }
}